I have been riding road bikes for decades with clipless pedals and on the flats try to clear the pedal on the backstroke rather than forcefully pulling up. It feels odd when I ride a bike with standard pedals as I am coming off the pedal on the back stroke. Regarding LeMond, the other technique he would discuss is, when climbing, to shift further back on the saddle (effectively increasing the distance from the pedals) and pull up, but at a lower cadence. The opposite is when sprinting, to move slightly forward on the saddle, maximize cadence and focus on the downstroke. The ‘shift in the saddle and change in cadence’ technique may negate the findings and conclusions of these studies.